The Ants of Africa
SUBFAMILY DORYLINAE - Subgenus Anomma comparative study and revision 2002, amended 2007
Subgenus Anomma introduction

This page deals with the sources of information on the subgenus Anomma and gives a suggested separation of the described species into species-groupings - a key to separate the species is linked at Subgenus Anomma key to species; also examined are the Subgenus Anomma comparison of the gastral apices.

Note - At the time of writing, December 2007, Caspar Schöning and Daniel Kronauer, are preparing a fresh analysis of Dorylus taxonomy. I anticipate that may show some of my interpretations of the species to be incorrect. In April (2008) Schöning et al. have synonymized D. gerstaeckeri and lamottei under Dorylus gribodoi, note they decline to separate it as in Anomma or Dorylus - http://www.mapress.com/zootaxa/2008/f/zt01749p052.pdf.


Species known only from sexual stages

Dorylus (Anomma) bishyiganus van Boven - queen
Dorylus (Anomma) funereus Emery - male; junior synonyms acherontus Santschi, pardus Santschi, stygis Santschi and zumpti Santschi
Dorylus (Anomma) niarembensis van Boven - queen
Dorylus (Anomma) nigricans Illiger - male (but see below)
Dorylus (Anomma) stanleyi Forel - male

The overall size, TL ca 19 mm, and mandible shape would appear to place Dorylus atratus Smith in subgenus Dorylus and not Anomma as given by Emery (1895j: 723).


I have tried to separate what is known of the males on MALES.


Time line of Anomma forms

Name First author Present status grouping & images
nigricans - male only; workers by later deduction Illiger, 1802 Dorylus nigricans yes - male & possible workers, Ghana & Schöning, Ivory Coast
burmeisteri - worker only Shuckard, 1840 Dorylus burmeisteri yes - Humle 13, Cameroon 11
arcens - worker only Westwood, 1847b Dorylus arcens yes, Cameroun, Benin & Guinea - Humle 07
rubellus - worker only Savage, 1849 Dorylus rubellus yes, Schöning CAR
molestus - worker only Gerstæcker, 1858 Dorylus molestus yes, Schöning Kenya
erraticus - minor worker only F Smith, 1865 Dorylus erraticus no merit
gribodoi - male only (as in Schoning et al, 2008) Emery, 1895j Dorylus gribodoi yes, as lamottei from Guinea
pubescens j. syn. of arcens Roger, 1861a; syn. Emery, 1895j, after examining type Dorylus arcens -
funereus - male only Emery, 1895j Dorylus funereus yes, male, CAR
gerstaeckeri - worker only - j. syn. of gribodoi Emery, 1895j Dorylus gribodoi yes, as lamottei from Guinea
emeryi - worker only Mayr, 1896 Dorylus emeryi yes, Schöning, Ivory Coast; drawing CRIN Nigeria
sjoestedti (as ssp of nigricans) - worker & queen Emery, 1899e Dorylus sjoestedti yes, CAR, Degallier; Schöning, Ivory Coast
wilverthi - all forms known Emery, 1899e Dorylus wilverthi yes, Uganda 0310103 (Schöning - Zaire)
pulsi ssp of emeryi - worker only Forel, 1904d Dorylus emeryi -
kohli - worker only Wasmann, 1904b Dorylus kohli yes, Guinea - Humle 11, Humle 2
opacus (as ssp of emeryi) - worker & male known Forel, 1909b Dorylus opacus yes, Schöning, Nigeria
stanleyi - male only Forel, 1909b Dorylus stanleyi -
congolensis (as variety of kohli) - worker only Santschi, 1910c Dorylus congolensis yes, Gabon - Pongara
minor variety of kohli Santschi, 1911g Dorylus kohli -
nigritarsis queen j syn of wilverthi Strand 1911 Dorylus wilverthi -
hybridus [hybrida] variety of burmeisteri Santschi, 1912b Dorylus burmeisteri -
ornatus (as variety of burmeisteri) - worker only Santschi, 1912b Dorylus ornatus yes, Uganda III-138 & Tanzania Mahale 05-03-02, 02-03-02, 08-03-02
pallidus variety of burmeisteri Santschi, 1912b Dorylus burmeisteri -
mayri - worker only Santschi, 1912b Dorylus mayri yes, Uganda III-135
quadratus (as variety of gerstaeckeri) - worker only Santschi, 1914a Dorylus quadratus yes, Tanzania, Mahale 03-03-02
rufescens (as stirps of nigricans) - worker & queen Santschi, 1915c - redefined from Wasmann, 1904b Dorylus rufescens yes, CAR, site GJ
frenisyi ssp of kohli Forel, 1916 Dorylus kohli -
sjoestedtiwilverthi j syn of wilverthi Wasmann, 1917 Dorylus wilverthi -
victoriae (as var. of kohli) - worker only Santschi, 1921c Dorylus victoriae yes, Ivory Coast - Humle 12; Guinea - Humle 3; Ghana 06G0066; Schöning, Ivory Coast labelled D. mayri
chapini ssp of kohli Wheeler, 1922 Dorylus kohli -
langi j syn of victoriae Wheeler, 1922 Dorylus victoriae -
militaris (as var. of kohli) - worker only Santschi, 1923e Dorylus militaris yes, Guinea - Humle 06, Humle 08
terrificus (as variety of nigricans) - worker only Santschi, 1923e Dorylus terrificus yes, Guinea - Humle 09
titan - worker only Santschi, 1923e Dorylus titan yes, Guinea - Humle 04
indocilis (as var. of kohli) - worker only Santschi, 1933b Dorylus indocilis yes, Tanzania - Mahale 28-02-02
nomadas queen j syn of wilverthi Santschi, 1933b Dorylus wilverthi -
vinalli var. of titan Santschi, 1933b Dorylus titan -
acherontus - variety of funereus - male only Santschi, 1937b Dorylus funereus -
pardus - variety of funereus - male only Santschi, 1937b Dorylus funereus -
stygis - variety of funereus - male only Santschi, 1937b Dorylus funereus -
zumpti - male only Santschi, 1937b Dorylus zumpti new status 2009
lamottei - worker only Bernard, 1953b Dorylus gribodoi yes, Guinea - Humle 05; Schöning, Ivory Coast CASENT0172626
bishyiganus - queen only van Boven, 1972 Dorylus bishyiganus no
niarembensis - queen only van Boven, 1972 Dorylus niarembensis no
ueharae - worker only new species - this website Dorylus ueharae yes, Tanzania - Mahale 27-02-02
Dorylus (Anomma) new species CAR IB - male only this site Dorylus (Anomma) new species CAR IB yes, Central African Republic

New key to species-groups 2007 - grouping derived from original descriptions and relationships

See also the Subgenus Anomma comparison of the gastral apices

. Major or soldier Minima Grouping
. Long-legged with petioles longer than they are wide .
1 {Dorylus burmeisteri face}Head widest mid-length, near parallel sided forward of that point, with rounded posterior angles; mandibles moderate; scapes relatively long & slender; in profile head depth ca 0.75 of HW and HL; TL max ca 12.5 mm

{Dorylus burmeisteri minima}

burmeisteri-group
burmeisteri, molestus, nigricans & rubellus
2 {Dorylus arcens face}Large, TL ca 13 mm; head widest at about the one-fifth point, with smoothly convex sides and sharp posterior angles; in profile head bulbous much deeper than other groups; mandibles very long and sharp; scapes relatively long & slender; generally more robust than burmeisteri-gp. Apart from the elongated processes at the postero-lateral extremities of the head wilverthi generally fits with the other arcens-group species {Dorylus arcens minima} arcens-group
arcens, mayri, terrificus & wilverthi
3 {Dorylus sjoestedti major}Characterised by the head being widest at the anterior end and narrowing significantly from front to back, with almost straight sides and sharp posterior angles; the latter never produced into distinct teeth (unlike wilverthi) {Dorylus sjoestedti minima} sjoestedti-group
rufescens, sjoestedti & victoriae
. Short-legged with petioles shorter than they are wide .
4 {Dorylus emery major}Relatively large, TL to 13 mm; head near square, rectangular; characterised by the short domed petiole, which seen from above is straight-sided and near square {Dorylus titan minima} emeryi-group
emeryi, opacus & titan
5 {Dorylus lamottei}Small less than 8 mm; head square, petiole short and domed/globular - possible link to Dorylus (Dorylus) {Dorylus lamottei minima} gribodoi-group
gribodoi, ueharae and Gabon species
6 {Dorylus congolensis major}Head near rectangular distinctly longer than wide and narrower posteriorly; petiole short in profile generally domed (some with a flat top); generally not more than TL 10 mm (congolensis to 12.5 mm) {Dorylus congolensis minima} kohli-group
congolensis, indocilis, kohli & militaris & quadratus

Contents Subgenus Anomma introduction
© 2007, 2008, 2009 - Brian Taylor CBiol FIBiol FRES
11, Grazingfield, Wilford, Nottingham, NG11 7FN, U.K.

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